[1] European Mol Biol Lab, Meyerhofstr 1, D-69117 Heidelberg, Germany
[2] Univ St Andrews, Scottish Oceans Inst, St Andrews KY16 8LB, Fife, Scotland
[3] Kitware Inc, 1712 Route 9,Suite 300, Clifton Pk, NY 12065 USA
[4] Scottish Assoc Marine Sci, Oban PA37 1QA, Argyll, Scotland