Taxonomic revision of the Phanaeus endymion species group (Coleoptera: Scarabaeidae), with the descriptions of five new species

The Phanaeus endymion species group is defined as a lineage of dung beetles distributed from Mexico to Ecuador. The current arrangement of the P endymion species group includes 18 species (five newly described and three revalidated herein): P arletteae Arnaud, 2018; P bravoensis Moctezuma, Sanchez-Huerta & Halffter, 2017; P chiapanecus sp. nov.; P edmondsi Moctezuma, Deloya & Halffter, 2019; P endymion Harold, 1863; P funereus Balthasar, 1939 stat. rev.; P halffierorum Edmonds, 1979; P huichol Moctezuma, Sanchez-Huerta & Halffter, 2017; P jackenioi sp. nov.; P malyi Arnaud, 2002; P olsoufieffi Balthasar, 1939 stat. rev.; P. pacificus sp. nov.; P. panamensis sp. nov.; P. porioni Arnaud, 2001 stat. rev.; P pyrois Bates, 1887; P. rzedowskii sp. nov.; P zapotecus Edmonds, 2006; and P zoque Moctezuma & Halffter, 2017. Phanaeus dionysius Kohlmann, Arriaga-Jimenez & Ros, 2018 syn. nov. is considered as a new junior subjective synonymy of P zapotecus Edmonds, 2006. Phanaeus blanchardi Olsoufieff, 1924 and P. bothrus Blackwelder, 1944 are junior objective synonyms of P. olsoufieffi Balthasar, 1939 stat. rev.