[1] Guangxi Zhuang Autonomous Reg, Guangxi Inst Bot, Guangxi Key Lab Plant Conservat & Restorat Ecol K, Guilin 541006, Peoples R China
[2] Chinese Acad Sci, Guilin 541006, Peoples R China
[3] South China Agr Univ, Coll Life Sci, Guangzhou 510642, Guangdong, Peoples R China
[4] Royal Bot Gardens, Identificat & Naming Dept, Richmond TW9 3AE, Surrey, England