[1] Southern Univ Sci & Technol, Sch Environm Sci & Engn, Shenzhen, Peoples R China
[2] Univ Copenhagen, Dept Geosci & Nat Resource Management, Copenhagen, Denmark
[3] Lund Univ, Dept Phys Geog & Ecosyst Sci, Lund, Sweden
[4] Univ Copenhagen, Terr Ecol Sect, Dept Biol, Copenhagen, Denmark