[1] Kyoto Univ, Grad Sch Sci, Human Evolut Studies Lab, Sakyo Ku, Kyoto 6068502, Japan
[2] Univ Tokyo, Grad Sch Frontier Sci, Dept Integrated Biosci, Kashiwanoha 5-1-5, Kashiwa, Chiba 277856, Japan
[3] St Marianna Univ, Dept Anat, Sch Med, Sugao 2-16-1, Kawasaki, Kanagawa 2168511, Japan
[4] Univ Tokyo, Grad Sch Sci, Dept Biol Sci, Bunkyo Ku, Hongo 7-3-1, Tokyo 1130033, Japan
[5] Univ Tokyo, Univ Museum, Bunkyo Ku, Hongo 7-3-1, Tokyo 1130033, Japan