[1] Univ Kent, Sch Anthropol & Conservat, Durrell Inst Conservat & Ecol DICE, Canterbury, Kent, England
[2] Univ Malaysia Sabah, Inst Trop Biol & Conservat, Kota Kinabalu, Sabah, Malaysia
[3] Forest Res Ctr Sepilok, Sabah Forestry Dept, Sandakan, Malaysia
[4] Imperial Coll London, Dept Life Sci, Silwood Pk, Ascot, Berks, England
[5] Univ Oxford, Dept Zool, Oxford, England
[6] Univ Cambridge, Dept Zool, Cambridge, England
[7] Univ Aberdeen, Sch Biol Sci, Aberdeen, Scotland
[8] Adam Mickiewicz Univ, Fac Biol, Evolutionary Biol Grp, Poznan, Poland
[9] Danum Valley Field Ctr, South East Asia Rainforest Res Partnership, Lahad Datu, Malaysia
[10] Queen Mary Univ London, Sch Biol & Chem Sci, London, England
[11] Natl Univ Singapore, Dept Biol Sci, Singapore, Singapore
[12] Nanyang Technol Univ, Asian Sch Environm, Singapore, Singapore