[1] Univ Fed Minas Gerais, Dept Genet Ecol & Evolut, Belo Horizonte, MG, Brazil
[2] Univ Estadual Paulista, UNESP, Dept Biol, Fac Ciencias Agr & Vet, Jaboticabal, SP, Brazil
[3] TU Bergakad Freiberg, Inst Biosci, Biol Ecol Unit, Freiberg, Germany
[4] Univ Fed Rio Grande do Sul, Dept Bot, Porto Alegre, RS, Brazil