[1] Swansea Univ, Dept BioSci, Singleton Pk, Swansea SA2 8PP, W Glam, Wales
[2] Alpine Wildlife Res Ctr, Parco Nazl Gran Paradiso,Degioz 11, I-11010 Valsavarenche, Aosta, Italy
[3] Univ Genoa, Dipartimento Sci Terra Ambiente & Vita, DISTAV, I-16132 Genoa, Italy
[4] Univ Trento, Dipartimento Ingn Civile Ambientale & Meccan, Via Mesiano 77, I-38123 Trento, Italy
[5] IASMA Res & Innovat Ctr, Fdn E Mach, Sustainable Ecosyst & Bioresources Dept, Via E Mach 1, I-38010 San Michele All Adige, Trento, Italy
[6] Univ Milan, Dipartimento Biosci, DBS, Via Celoria 26, I-20133 Milan, Italy
[7] Univ Toulouse, CNRS, UPS, CRCA,CBI, Bat 4R3,118 Route Narbonne, F-31062 Toulouse 9, France
[8] Univ Pavia, DSTA, Via Ferrata 9, I-27100 Pavia, Italy