[1] Univ Tennessee, Ctr Hlth Sci, Ophthalmol, Memphis, TN 38163 USA
[2] Univ Tennessee, Ctr Hlth Sci, Genet Genom & Informat, Memphis, TN 38163 USA
[3] Univ Tennessee, Ctr Hlth Sci, Pharmacol, Memphis, TN 38163 USA
[4] Jackson Lab, 600 Main St, Bar Harbor, ME 04609 USA
[5] Univ Utah, Sch Med, Ophthalmol, Salt Lake City, UT USA
[6] Emory Univ, Ophthalmol, Atlanta, GA 30322 USA