[1] Capital Normal Univ, Coll Resource Environm & Tourism, Beijing 100048, Peoples R China
[2] Helmholtz Ctr Polar & Marine Res, Res Unit Potsdam, Alfred Wegener Inst, D-14473 Potsdam, Germany
[3] Univ Potsdam, Inst Environm Sci & Geog, D-14476 Potsdam, Germany
[4] Univ Potsdam, Inst Biochem & Biol, D-14476 Potsdam, Germany
[5] Chinese Acad Sci, Inst Tibetan Plateau Res, Key Lab Alpine Ecol, Alpine Paleoecol & Human Adaptat Grp ALPHA, Beijing 100101, Peoples R China
[6] Chinese Acad Sci, CAS Ctr Excellence Tibetan Plateau Earth Sci, Beijing 100101, Peoples R China
[7] Zhejiang Normal Univ, Coll Chem & Life Sci, Jinhua 321004, Zhejiang, Peoples R China
[8] Chinese Acad Sci, Inst Geog Sci & Nat Resources Res, Beijing 100101, Peoples R China