[1] Chinese Acad Sci, South China Sea Inst Oceanol, CAS Key Lab Trop Marine Bioresources & Ecol, Guangzhou 510301, Peoples R China
[2] Southern Marine Sci & Engn Guangdong Lab Guangzhou, Guangzhou 511458, Peoples R China
[3] Guangdong Prov Key Lab Marine Resources & Coastal, Guangzhou 510006, Peoples R China
[4] Minist Nat Resources, South China Sea Bur, South China Sea Environm Monitoring Ctr, Guangzhou 510300, Peoples R China
[5] Univ Chinese Acad Sci, Beijing 100049, Peoples R China
[6] Chinese Acad Sci, South China Sea Inst Oceanol, Guangzhou 510301, Peoples R China