[1] Univ Vet Med Hannover, Inst Terr & Aquat Wildlife Res, D-25761 Busum, Schleswig Holst, Germany
[2] Univ Liege, Freshwater & Ocean Sci Unit Res Oceanol, Allee 6 Aout,B6C, B-4000 Liege, Belgium
[3] Univ Potsdam, Inst Biochem & Biol, Unit Evolutionary Biol Systemat Zool, Karl Liebknecht Str 24-25, D-14476 Potsdam, Germany
[4] Univ Utrecht, Fac Vet Med, Dept Pathobiol, Yalelaan 1, NL-3584 CL Utrecht, Netherlands
[5] Ctr Environm, Fisheries & Aquaculture Sci Cefas Lowestoft Lab, Pakefield Rd, Lowestoft NR33 0HT, Suffolk, England
[6] Inst Zool, CSIP, Regents Pk, London NW1 4RY, England
[7] SRUC Wildlife Unit, Inverness IV2 4JZ, Scotland
[8] Univ Liege, CART, LEAE, B6c, Liege, Belgium