[1] Louisiana State Univ, Museum Nat Sci, Baton Rouge, LA 70803 USA
[2] Louisiana State Univ, Dept Biol Sci, Baton Rouge, LA 70803 USA
[3] Univ Maryland, Behav Ecol Evolut Systemat Program, College Pk, MD 20742 USA
[4] Museu Paraense Emilio Goeldi, BR-66040170 Belem, Para, Brazil
[5] Smithsonian Inst, USGS Patuxent Wildlife Res Ctr, Natl Museum Nat Hist, Washington, DC 20013 USA