[1] Univ Guadalajara, Dept Bot & Zool, Ctr Univ Ciencias Biol & Agr, Camino Ing Ramon Padilla Sanchez 2100, Zapopan 45200, Jalisco, Mexico
[2] Univ Guadalajara, Lab Nacl Identificat & Caracterizac Vegetal LaniV, Ctr Univ Ciencias Biol & Agr, Camino Ing Ramon Padilla Sanchez 2100, Zapopan 45200, Jalisco, Mexico
[3] Inst Ecol AC, Red Biol Evolut, Carretera Antigua Coatepec 351, Xalapa 91073, Veracruz, Mexico
[4] Iowa State Univ, Dept Ecol Evolut Otganismal Biol, Ames, IA 50011 USA