[1] Univ Lille 2, Inst Reg Rech Agroalimentaire & Biotechnol Charle, EA1026, F-59000 Lille, France
[2] Ctr Biotechnol Technopole Borj Cedria CBBC, Lab Plantes Extremophiles, Hammam Lif 2050, Tunisia
[3] INSERM, U995, UFR Pharm, UDSL, F-59000 Lille, France
[4] Catholic Univ Louvain, Pharmacognosy Res Grp, Louvain Drug Res Inst, B-1200 Brussels, Belgium
[5] Univ Lille Nord France, Inst Pasteur Lille, INSERM, U1019,CNRS,UMR8204, Lille, France