[1] South African Natl Biodivers Inst, Biosystemat Res & Biodivers Collect Div, ZA-0001 Pretoria, South Africa
[2] Univ Pretoria, Dept Plant Sci, HGWJ Schweickerdt Herbarium, ZA-0002 Pretoria, South Africa
[3] Univ Coimbra, Dept Ciencias Vida, Ctr Funct Ecol, P-3001455 Coimbra, Portugal
[4] Nelson Mandela Metropolitan Univ, Dept Bot, ZA-6031 Port Elizabeth, South Africa
[5] South African Natl Biodivers Inst, Ethnobot Unit, ZA-0001 Pretoria, South Africa
[6] Univ KwaZulu Natal, Sch Chem & Phys, ZA-4041 Durban, South Africa
[7] Utah State Univ, Intermt Herbarium, Logan, UT 84322 USA