[1] Sao Paulo State Univ, Aquaculture Ctr, Jaboticabal, SP, Brazil
[2] SAA SP, Fishery Inst, Sao Paulo, SP, Brazil
[3] Univ South & Southern Para Unifesspa, Biol Fac FACBIO, Grp Studies Reprod Amazon Fish GERPA LANEC, Maraba, Para, Brazil
[4] Chico Mendes Inst Biodivers Conservat, Natl Ctr Res & Conservat Continental Fish, Pirassununga, SP, Brazil
[5] Hokkaido Univ, Fac Fisheries Sci, Hakodate, Hokkaido, Japan