[1] Univ Witwatersrand, Sydney Brenner Inst Mol Biosci, Fac Hlth Sci, Johannesburg, South Africa
[2] Maastricht Univ, Med Ctr, Dept Clin Genet, Maastricht, Netherlands
[3] Maastricht Univ, Med Ctr, GROW Sch Oncol & Dev Biol, Maastricht, Netherlands
[4] Radboud Univ Nijmegen, Med Ctr, Dept Human Genet, Donders Inst Brain Cognit & Behav, Nijmegen, Netherlands
[5] Univ Edinburgh, Ctr Trop Livestock Genet & Hlth, Edinburgh, Midlothian, Scotland