[1] Univ Wurzburg, Bioctr, D-97074 Wurzburg, Germany
[2] Keio Univ, Sch Med, Dept Mol Biol, Tokyo 1608582, Japan
[3] European Mol Biol Lab, Dev Biol Programme, D-69012 Heidelberg, Germany
[4] ERATO, Sakyo Ku, Kyoto 6068305, Japan
[5] Max Planck Inst Mol Genet, D-14195 Berlin, Germany
[6] Natl Univ Singapore, Dept Biol Sci, Singapore 117543, Singapore
[7] Nagoya Univ, Div Biol Sci, Grad Sch Sci, Nagoya, Aichi 4648602, Japan
[8] Univ Tokyo, Dept Biol Sci, Grad Sch Sci, Bunkyo Ku, Tokyo 1130033, Japan
[9] Univ Tokyo, Dept Integrated Biosci, Grad Sch Frontier Sci, Kashiwa, Chiba 2778562, Japan