[1] Pontificia Univ Catolica Rio Grande do Sul, Lab Vertebrate Systemat, Ave Ipiranga 6681, BR-90619900 Porto Alegre, RS, Brazil
[2] Univ Louisiana Lafayette, Lafayette, LA 70504 USA
[3] Univ Fed Rio de Janeiro UFRJ, Nucleo Ecol & Desenvolvimento Socioambiental Maca, Grp Sistemat & Biol Evolut, Caixa Postal 119331, BR-27910970 Macae, RJ, Brazil
[4] Harvard Univ, Museum Comparat Zool, 26 Oxford St, Cambridge, MA 02138 USA
[5] Calif Acad Sci, Nat Conservancy & Sect Ichthyol, 55 Mus Concourse Dr, San Francisco, CA 94118 USA