[1] Univ Utrecht, Fac Geosci, Copernicus Inst Sustainable Dev, Princetonlaan 8a, NL-3584 CB Utrecht, Netherlands
[2] Univ Utrecht, Inst Environm Biol, Sect Ecol & Biodivers, Padualaan 8, NL-3584 CH Utrecht, Netherlands
[3] Tropenbos Int, Lawickse Allee 11, NL-6701 AN Wageningen, Netherlands
[4] Lund Univ, Ctr Environm & Climate Res, Solvegatan 37, S-22362 Lund, Sweden
[5] Univ Zurich, Univ Res Prior Program Global Change & Biodivers, Winterthurerstr 190, Zurich, Switzerland
[6] Univ Zurich, Dept Geog, Winterthurerstr 190, Zurich, Switzerland