[1] Royal Tyrrell Museum Palaeontol, POB 7500, Drumheller, AB T0J 0Y0, Canada
[2] Univ Lethbridge, Dept Geog, 4401 Univ Dr West, Lethbridge, AB T1K 3M4, Canada
[3] Univ Lethbridge, Dept Math & Comp Sci, 4401 Univ Dr West, Lethbridge, AB T1K 3M4, Canada
[4] Univ Toronto, Dept Ecol & Evolutionary Biol, 25 Willcocks St, Toronto, ON M5S 3B2, Canada
[5] Okayama Univ Sci, Dept Biosphere Geosphere Sci, Kita Ku, 1-1 Ridaicho, Okayama, Okayama 7000005, Japan