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Wonkwang Univ, Fac Biol Sci, 460 Iksan Daero, Iksan Jeonbuk 54538, South Korea
Wonkwang Univ, Inst Basic Sci, 460 Iksan Daero, Iksan Jeonbuk 54538, South Korea Incheon Natl Univ, Dept Marine Sci, 119 Acad Ro, Incheon 22012, South Korea