A NEW TRISTICHOPTERID (SARCOPTERYGII, TETRAPODOMORPHA) FROM THE UPPER FAMENNIAN EVIEUX FORMATION (UPPER DEVONIAN) OF BELGIUM

Additional material of a large specimen of tristichopterid fish from the Upper Famennian Evieux Formation of Belgium is described. This large fish was previously assigned to Tristichopteridae gen. et sp. indet. due to the lack of diagnostic anatomical data. New available material consists of the internal surface of the parietal shield, vomers and anterior part of the parasphenoid, subopercular and submandibulo-branchiostegal bones, and an internal view of the anterior part of the mandible. A possible autapomorphy of the new form from Belgium, Langlieria socqueti gen. nov. et sp. nov., is the absence of marginal teeth on the vomer except on its most lateral part. Apart from these features, it only differs from the genus Mandageria from Australia in the absence of marginal teeth between the dentary fang and the mandibular symphysis, in the presence of a raised marginal crest lateral to the anterior coronoid fang, and in the presence of numerous small marginal teeth on the premaxilla. It differs from the cosmopolitan genus Eusthenodon in a number of respects: the supratemporal, tabular, and postparietal bones are superficially fused, as are the intertemporal and parietal bones, the dermal ornament is proportionally very fine, and the denticulated field of the parasphenoid stands proud rather than being recessed into the body of the bone.