[1] Univ Texas Austin, Dept Mol Biosci, Austin, TX 78712 USA
[2] Univ Cambridge, Dept Phys, Cavendish Lab, JJ Thomson Ave, Cambridge CB3 0HE, England
[3] Univ Cambridge, Canc Res UK Gurdon Inst, Wellcome Trust, Tennis Court Rd, Cambridge CB2 1QN, England
[4] Univ New South Wales, Sch Med Sci, Single Mol Sci node, EMBL Australia, Sydney, NSW 2052, Australia
[5] Univ Sydney, Sydney Med Sch, Centenary Inst, Sydney, NSW 2006, Australia