[1] Univ Birmingham, Inst Canc & Genom Sci, Birmingham, W Midlands, England
[2] Univ Hosp Birmingham, Dept Gastroenterol, Birmingham, W Midlands, England
[3] Univ Birmingham, Inst Microbiol & Infect, Birmingham, W Midlands, England
[4] Univ Birmingham, Liver Res Ctr, Birmingham, W Midlands, England
[5] Univ Birmingham, Natl Inst Hlth Res NIHR Birmingham, Liver Biomed Res Unit BRU, Inst Immunol & Immunotherapy, Birmingham, W Midlands, England
[6] Univ Oslo, Oslo Univ Hosp, Rikshosp, Norwegian PSC Res Ctr, Oslo, Norway
[7] Inst Clin Med, Oslo, Norway
[8] Kings Coll London, Twin Res & Genet Epidemiol, London, England
[9] Univ Birmingham, Inst Immunol & Immunotherapy, Birmingham, W Midlands, England
[10] Univ Hosp Birmingham, Inst Translat Med, Birmingham, W Midlands, England