[1] Korea Inst Ocean Sci & Technol, East Sea Environm Res Div, Gyeongbuk, South Korea
[2] Univ Sci & Technol, Dept Marine Biotechnol, Taejon, South Korea
[3] Natl Fisheries Res & Dev Inst, Tidal Flat Res Inst, Kunsan, South Korea
[4] Rural Dev Adm, Natl Inst Anim Sci, Chungnam, South Korea
[5] Univ Sci & Technol, Dept Marine Biol, Taejon, South Korea
[6] Korea Inst Ocean Sci & Technol, Marine Ecosyst Res Div, Ansan 426744, South Korea