[1] Inst Rech Dev, UMR EME 212, 16 Rue Claude Chappe, F-97420 Le Port, Reunion, France
[2] Oceans & Coasts Res, Dept Environm Affairs, ZA-8000 Cape Town, South Africa
[3] Bayworld Ctr Res & Educ, ZA-8012 Cape Town, South Africa
[4] Univ Cape Town, Marine Res Inst, Nansen Tutu Ctr Arine Environm Res, ZA-7701 Rondebosch, South Africa
[5] Nansen Environm & Remote Sensing Ctr, Bergen, Norway
[6] Rhodes Univ, ZA-6140 Grahamstown, South Africa