[1] Inst Tecnol Vale, Rua Boaventura da Silva 955, BR-66055090 Belem, Para, Brazil
[2] Museu Paraense Emilio Goeldi, Coord Bot, Av Perimetral 1901, BR-66077830 Belem, Para, Brazil
[3] Univ Fed Vales Jequitinhonha & Mucuri, Dept Ciencias Biol, Campus JK,Rodovia MG 367,5-000, BR-39100000 Diamantina, MG, Brazil
[4] Field Museum Nat Hist, Keller Sci Act Ctr, 1400 S Lakeshore Dr, Chicago, IL 60605 USA
[5] Univ Sao Paulo, Inst Biociencias, Dept Bot, Lab Sistemat Vegetal, Rua Matao 277, BR-05508900 Sao Paulo, Brazil