[1] James Hutton Inst, Dundee DD2 5DA, Scotland
[2] Univ Hull, Dept Biol Sci, Kingston Upon Hull HU6 7RX, N Humberside, England
[3] Zool Soc London, Inst Zool, London NW1 4RY, England
[4] W Virginia Univ, Div Forestry & Nat Resources, Morgantown, WV 26506 USA
[5] Univ Missouri, Sch Nat Resources, Columbia, MO USA
[6] Univ Cape Town, Dept Stat Sci, ZA-7700 Rondebosch, South Africa
[7] Univ Washington, Sch Aquat & Fishery Sci, Seattle, WA 98195 USA
[8] Univ N Texas, Inst Appl Sci, Inst Appl Sci, Denton, TX 76203 USA