[1] MRC, Anat Neuropharmacol Unit, Oxford OX1 3TH, England
[2] Univ Leeds, Sch Biomed Sci, Leeds LS2 9JT, W Yorkshire, England
[3] Univ London Imperial Coll Sci Technol & Med, Sch Med, London SW7 2AZ, England
[4] Univ Birmingham, Sch Med, Div Neurosci, Birmingham B15 2TT, W Midlands, England