[1] Russian Acad Sci, Inst Physicochem & Biol Problems Soil Sci, Pushchino 142290, Russia
[2] Helmholtz Ctr Polar & Marine Res, Alfred Wegener Inst, D-14401 Potsdam, Germany
[3] Lomonosov Moscow State Univ, Fac Geog, Lab Geoecol North, Moscow 119991, Russia
[4] Univ Potsdam, Inst Geosci, D-14469 Potsdam, Germany