[1] CNRS, Moulis, France
[2] Univ Paul Sabatier Toulouse III, SETE, UMR 5321, Moulis, France
[3] Univ Paul Sabatier Toulouse III, UMR 5174, Lab Evolut & Diversite Biol, Toulouse, France
[4] Eawag Swiss Fed Inst Aquat Sci & Technol, Dept Aquat Ecol, Dubendorf, Switzerland
[5] Univ Zurich, Dept Evolutionary Biol & Environm Studies, Zurich, Switzerland
[6] Univ Florida, Dept Biol, Gainesville, FL USA
[7] Univ Montpellier 2, CNRS, IRD, Inst Sci Evolut,EPHE,CC65, Montpellier, France
[8] Catholic Univ Louvain, Earth & Life Inst, Biodivers Res Ctr, Louvain La Neuve, Belgium
[9] Univ Aberdeen, Sch Biol Sci, Aberdeen, Scotland