[1] No Arizona Univ, Sch Forestry, Bat Ecol & Genet Lab, Flagstaff, AZ 86011 USA
[2] No Arizona Univ, Pathogen & Microbiome Inst, Flagstaff, AZ 86011 USA
[3] Univ Adelaide, Dept Anim & Vet Sci, Waite Campus, Glen Osmond, SA 5064, Australia
[4] Univ Adelaide, Sch Publ Hlth, Adelaide, SA 5005, Australia