[1] Univ Cambridge, Dept Geog, Cambridge, England
[2] Univ Ghent, Dept Biol, Limnol Unit, Ghent, Belgium
[3] Univ Ghent, Dept Biol, Renard Ctr Marine Geol, Ghent, Belgium
[4] Univ Lancaster, Lancaster Environm Ctr, Lancaster, England
[5] Trinity Coll Dublin, Dept Geol, Dublin, Ireland