[1] Univ Manchester, Ctr Musculoskeletal Res, Manchester 2, Lancs, England
[2] Univ Manchester, Dermatol Ctr, Manchester, Lancs, England
[3] Univ Leeds, Leeds, W Yorkshire, England
[4] Univ Coll Dublin, Dublin, Ireland
[5] Newcastle Univ, Inst Cellular Med, Newcastle Upon Tyne, Tyne & Wear, England
[6] Manchester Acad Hlth Sci Ctr, NIHR Manchester Musculoskeletal Biomed Res Unit, Manchester, Lancs, England