[1] Hunan Univ Sci & Technol, Hunan Prov Key Lab Econ Crops Genet Improvement &, Xiangtan 411201, Peoples R China
[2] Chinese Acad Sci, Inst Zool, Key Lab Zool Systemat & Evolut, Beijing 100101, Peoples R China
[3] China Agr Univ, Dept Entomol, Beijing 100193, Peoples R China
[4] Capital Normal Univ, Coll Life Sci, Beijing 100048, Peoples R China
[5] Capital Normal Univ, Acad Multidisciplinary Studies, Beijing 100048, Peoples R China