[1] Fed Univ Rio Grande do Norte UFRN, Biosci Ctr, Dept Bot & Zool, Lab Collembola, Highway BR-101,Campus Univ, BR-59072970 Natal, RN, Brazil
[2] Fed Inst Educ Sci & Technol Piaui, Biodivers & Biotechnol Res Grp, North Ctr Piaui, Pedro 2, Piaui, Brazil
[3] CPEN, Natl Inst Amazonian Res INPA, Lab Soil Invertebrate Systemat & Ecol, Manaus, Amazonas, Brazil