[1] Univ Edinburgh, Usher Inst Populat Hlth Sci & Informat, Edinburgh, Midlothian, Scotland
[2] Univ Tartu, Estonian Genome Ctr, Tartu, Estonia
[3] Broad Inst, Cambridge, MA USA
[4] Harvard Med Sch, Dept Genet, Boston, MA USA
[5] Univ Edinburgh, MRC Human Genet Unit, Edinburgh, Midlothian, Scotland