[1] Ocean Univ China, Coll Environm Sci & Engn, Qingdao, Peoples R China
[2] Minist Nat Resources, Inst Oceanog 1, Key Lab Marine Ecoenvironm Sci & Technol, Qingdao, Peoples R China
[3] Pilot Natl Lab Marine Sci & Technol Qingdao, Lab Marine Ecol & Environm Sci, Qingdao, Peoples R China