[1] Novartis Inst BioMed Res, CH-4002 Basel, Switzerland
[2] Univ Tubingen, Sect Neurobiol Eye, D-72076 Tubingen, Germany
[3] Queensland Univ Technol, Sch Optometry, Brisbane, Qld 4001, Australia
[4] Queensland Univ Technol, Inst Hlth & Biomed Innovat, Brisbane, Qld 4001, Australia