[1] Univ Concepcion, Interdisciplinary Ctr Aquaculture Res, OHiggins 1695, Concepcion, Chile
[2] Univ Concepcion, Dept Oceanog, Doctorate Program Oceanog, Casilla 160-C, Concepcion, Chile
[3] UPMC CNRS IRD MNHN, Lab Oceanog & Climatol Experimentat & Anal Numer, Inst Pierre Simon Laplace, 4 Pl Jussieu,Case 100, F-75252 Paris 05, France
[4] Univ Concepcion, Dept Geophys, Concepcion, Chile