[1] Forestry & Forest Prod Res Inst, Tsukuba, Ibaraki 3058687, Japan
[2] Gifu Univ, River Basin Res Ctr, Gifu 5011193, Japan
[3] Japan Int Res Ctr Agr Sci, Tsukuba, Ibaraki 3058686, Japan
[4] Natl Inst Humanities, Res Inst Humanity & Nat, Kita Ku, Kyoto 6038047, Japan