[1] Smithsonian Inst, Natl Museum Nat Hist, Washington, DC 20560 USA
[2] Scottish Marine Biol Assoc, Oban PA34 4AD, Argyll, Scotland
[3] USSR Acad Sci, Inst Oceanol, Moscow, Russia
[4] Russian Acad Sci, Inst Oceanol, Moscow 117218, Russia
[5] Russian Acad Sci, Inst Zool, St Petersburg 199034, Russia