[1] East China Normal Univ, Sch Life Sci, Dept Biol, Bryol Lab, 3663 Zhong Shan North Rd, Shanghai 200062, Peoples R China
[2] Hainan Univ, Res Ctr Terr Biodivers South China Sea, Inst Trop Agr & Forestry, 58 Renmin Rd, Haikou 570228, Hainan, Peoples R China
[3] Prince Songkla Univ, Fac Sci, Dept Biol, Hat Yai 90112, Songkhla, Thailand
[4] Royal Bot Gardens & Domain Trust, Natl Herbarium New South Wales, Mrs Macquaries Rd, Sydney, NSW 2000, Australia
[5] East China Normal Univ, Tiantong Natl Stn Forest Ecosyst, Shanghai Key Lab Urban Ecol Proc & Ecorestorat, 500 Dongchuan Rd, Shanghai 200241, Peoples R China