[1] Univ Calicut, Insect Ecol & Ethol Lab, Dept Zool, Calicut 673635, Kerala, India
[2] Russian Acad Sci, Zool Inst, St Petersburg 199034, Russia
[3] Polish Acad Sci, Museum & Inst Zool, Wilcza 64, Warsaw, Poland
[4] Ashoka Trust Res Ecol & Environm, Bangalore 560064, Karnataka, India