[1] Natl Sun Yat Sen Univ, Dept Oceanog, Kaohsiung 804, Taiwan
[2] State Ocean Adm, Inst Oceanog 2, Key Lab Marine Ecosyst & Biogeochem, Hangzhou 310012, Zhejiang, Peoples R China
[3] Natl Taiwan Ocean Univ, Inst Marine Environm & Ecol, Keelung 202, Taiwan
[4] ROC Naval Acad, Dept Appl Sci, Kaohsiung 813, Taiwan
[5] Natl Cheng Kung Univ, Inst Ocean Technol & Marine Affair, Tainan 701, Taiwan
[6] Natl Taiwan Ocean Univ, Dept Marine Environm Informat, Keelung 202, Taiwan