[1] James Cook Univ, Sch Earth & Environm Sci, Townsville, Qld 4811, Australia
[2] CSIRO Marine & Atmospher Res, CSIRO Wealth Oceans Natl Res Flagship, Hobart, Tas 7001, Australia
[3] James Cook Univ, Fishing & Fisheries Res Ctr, Townsville, Qld 4811, Australia