[1] Chinese Acad Sci, Inst Bot, State Key Lab Systemat & Evolutionary Bot, 20 Nanxincun, Beijing 100093, Peoples R China
[2] Univ Gottingen, Dept Palynol & Climate Dynam, Untere Karspule 2, D-37073 Gottingen, Germany
[3] Univ Vienna, Dept Palaeontol, Althanstr 14, A-1090 Vienna, Austria
[4] Chinese Acad Sci, Inst Earth Environm, SKLLQG, Xian 710075, Peoples R China
[5] CAS Ctr Excellence Tibetan Plateau Earth Sci, Beijing 100101, Peoples R China
[6] Univ Chinese Acad Sci, Coll Earth & Planetary Sci, Beijing 100049, Peoples R China
[7] Univ Chinese Acad Sci, Beijing 100049, Peoples R China