[1] Univ Georgia, Coverdell Ctr Biomed & Hlth Sci, Dept Infect Dis, 500 DW Brooks Dr, Athens, GA 30602 USA
[2] Univ Georgia, Grad Program Microbiol, Athens, GA 30602 USA
[3] Univ Georgia, Grad Program Infect Dis, Athens, GA 30602 USA
[4] Univ Georgia, Coll Vet Med, Ctr Vaccines & Immunol, 501 DW Brooks Dr, Athens, GA 30602 USA
[5] Emory Univ, Dept Biol, 1510 Clifton Rd, Atlanta, GA 30322 USA